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Exploring the beautiful nature of California
California Nature: Crustaceans
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There are least 77 species of crustaceans
that are considered nonindigenous to the waters in which they
occur. Out of these 77 species, 44 have become established in
their new environment. Introduction of at least one species has
occurred in the inland or coastal waters of 43 states.
California ranks the highest with over 50 species of
nonindigenous crustaceans, mostly in coastal waters and bays.
Crustaceans are a type of Arthropod. The name may not sound familiar, but you
probably know them. You may even have eaten one. Crustaceans have a hard,
external shell which protects their body. Crustaceans have a
head and abdomen. The head has antennae which are part of their
sensory system. The abdomen includes the heart, digestive system
and reproductive system.
The abdomen also has appendages, such as legs, for crawling and
swimming. Many crustaceans also have claws that help with
crawling and eating. The most commonly known California
crustaceans are the shrimp, lobster and crabs. |
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It is convenient to divide crustaceans into two groups, the
entomostracans and the malacostraceans. The entomostracans
include the smaller species, such as fairy shrimp, water fleas,
copepods, and barnacles. The malacostaceans include the larger,
more familiar crustaceans, such as crabs, shrimp, and lobsters.
All crustaceans have an exoskeleton made of the protein chitin and calcium. This
external shell, in addition to being protective, gives rigid support for the
attachment of the muscles. The exoskeleton is made of separate plates connected
by thin membranes. This segmented exoskeleton creates joints, allowing the
crustacean to move its body and appendages. Typically, the body of crustaceans is divided into two sections: the head and
the trunk. The head has five pairs of appendages: two pairs of antennae and
three feeding appendages, composed of one pair of mandibles and two pairs of
maxillae. The trunk is usually divided into the thorax and the abdomen. In many
crustaceans a shell called the carapace covers the head and thorax. Crustaceans
respirate via gills, located where the legs attach to the thorax.
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Most crustaceans are free-living, but some
are sessile (attached) and a few are even parasitic. Some filter
tiny plankton or even bacteria from the water, others are active
predators, while still others scavenge nutrients from detritus.
Although most crustaceans are dioecious (separate male and
female individuals), the actual mechanisms by which they
fertilize vary greatly. Some crustaceans hatch young that
resemble miniature adults, others go through larval stages.
Although a crustacean grows, its exoskeleton does not, so the
animal must molt its old exoskeleton in order to accommodate its
expanding body. The process of molting is hormonally controlled.
In preparation for molting, the tissue layer under the
exoskeleton detaches and secretes a new exoskeleton. At this
point, the animal has two skeletons – the outer one and the
inner one. When the new skeleton is completely formed, the old
skeleton splits along specific weak points and the animal pulls
out, leaving its old skeleton intact except for the split.
Mobility is limited immediately after a molt because the
exoskeleton is not rigid enough to keep the limbs stiff.
Although crustaceans molt throughout their entire life, they
molt less frequently with age. |
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Crustaceans are of great direct and indirect importance to
humans. The larger crustaceans (shrimps, lobsters, and crabs)
are used as food throughout the world, and are therefore
important to human economies. Small planktonic crustaceans, such
as copepods, water fleas, and krill, are a major link in the
marine food chain between the photosynthetic phytoplankton and
larger carnivores such as fish and whales. Still others are
crucial in recycling nutrients trapped in the bodies of dead
organisms.
Lobsters are heavy-bodied
decapods and are generally inhabitants of holes and crevices of
rocky bottoms. Using their four sets of paired legs, lobsters
walk along the bottom, feeling the path ahead by waving their
antennae. Their two front legs are actually claws – a large
crusher claw, and a smaller claw for transferring food to the
mouth. They also have two pairs of antennae (or feelers) and a
tail with broad, scaly plates on the end. They can swim by
flipping their fanlike tail forward violently.
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If you aren't yet convinced how important
crustaceans are, think about this: Crustaceamorpha are the
dominant arthropods, and very often the most numerically
abundant type of animal in aquatic environments. They make up a
significant portion of animal communities in all aquatic
habitats, from the most extreme (glacial ice or temporary desert
ponds) to the largest (deep ocean) and everywhere in
between. The Crustaceamorpha are arguably the most well known
of the arthropods because of their contributions to aquatic,
aerial, and terrestrial food webs.
Crustaceans are the arthropods that dominate marine habitats, but they are also found in large numbers in freshwater and a few groups have made their way successfully onto on land. When found on land crustaceans are either found in moist protected habitats like under logs or in leaf litter in cool forests, or they are encysted (enclosed in a tough protective capsule, nearly dried out, and dormant). |
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Crabs
Lobster Shrimp |
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